Unlock rules before spending points
The normal skill-tree purchases cost one skill point each. Leveling supplies points, but leveling alone doesn't bypass story progression. The skill menu opens after the introductory mission, Hold Onto Your Web-Shooters, and Camouflage becomes available through the story after Time to Rally.
The levels below are the listed level requirements, not promises that a power will work before its story introduction. Venom Dash Launcher, for example, improves Venom Dash; it doesn't replace the story unlock for Dash itself.
The nine Challenge skills are different. Complete the relevant Spider-Training challenge and the reward is yours without spending a skill point. Finish each category's 1.0 challenge to progress to 2.0, then 3.0. You do not need an Ultimate score to earn those skills.
Three final tree skills need New Game+: Let's Go! at level 21, Bio-Electric Instincts at 22 and Unseen Force at 23. So if three boxes remain locked on a completed first run, you haven't necessarily missed anything.
Unlock rules before spending points
· Games / Marvel's Spider-Man: Miles Morales